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NARMS taxon details

Doris Linnaeus, 1758

137914 (urn:lsid:marinespecies.org:taxname:137914)

  1. Biota
  2. Animalia
  3. Mollusca
  4. Gastropoda
  5. Heterobranchia
  6. Euthyneura
  7. Ringipleura
  8. Nudipleura
  9. Doridida
  10. Doridoidei
  11. Doridoidea
  12. Dorididae
  13. Doris
accepted
Genus
Doris verrucosa Linnaeus, 1758 (type by monotypy)
Dorididae Rafinesque, 1815
Dorididae Rafinesque, 1815
Doridigitata A. d'Orbigny, 1839 · unaccepted (junior objective synonym of...)  
junior objective synonym of Doris
Doris (Doridigitata) A. d'Orbigny, 1839 · unaccepted > junior objective synonym
Staurodoris Bergh, 1878 · unaccepted (Invalid: junior objective synonym...)  
Invalid: junior objective synonym of Doris; Placed on the Official Index by ICZN Opinion 1980

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  1. Species Doris alboranica Bouchet, 1977
  2. Species Doris berghi (Vayssičre, 1901)
  3. Species Doris bertheloti A. d'Orbigny, 1839
  4. Species Doris bicolor (Bergh, 1884)
  5. Species Doris januarii (Bergh, 1878)
  6. Species Doris kyolis (Ev. Marcus & Er. Marcus, 1967)
  7. Species Doris marmorata Risso, 1818
  8. Species Doris nobilis (Odhner, 1907)
  9. Species Doris ocelligera (Bergh, 1881)
  10. Species Doris pseudoverrucosa (Ihering, 1886)
  11. Species Doris sticta (Iredale & O'Donoghue, 1923)
  12. Species Doris verrucosa Linnaeus, 1758
  13. Species Doris flammea Alder & Hancock, 1844 (uncertain > nomen dubium)
  14. Subgenus Doris (Adalaria) Bergh, 1878 accepted as Adalaria Bergh, 1878
    1. Species Doris (Adalaria) sibirica Aurivillius, 1887 accepted as Calycidoris guentheri Abraham, 1876 (synonym)
  15. Subgenus Doris (Doridigitata) A. d'Orbigny, 1839 accepted as Doris Linnaeus, 1758 (unaccepted > junior objective synonym)
    1. Species Doris (Doridigitata) bertheloti A. d'Orbigny, 1839 accepted as Doris bertheloti A. d'Orbigny, 1839
  16. Species Doris affinis Gmelin, 1791 accepted as Flabellina affinis (Gmelin, 1791) (original combination)
  17. Species Doris albescens R. A. Philippi, 1836 accepted as Felimida purpurea (Risso, 1831) accepted as Rudmania purpurea (Risso, 1831)
  18. Species Doris arborescens O. F. Müller, 1776 accepted as Dendronotus frondosus (Ascanius, 1774)
  19. Species Doris areolata Stuwitz, 1836 accepted as Doris pseudoargus Rapp, 1827 accepted as Archidoris pseudoargus (Rapp, 1827) (synonym)
  20. Species Doris argo Linnaeus, 1767 accepted as Platydoris argo (Linnaeus, 1767)
  21. Species Doris aspera Alder & Hancock, 1842 accepted as Onchidoris muricata (O. F. Müller, 1776)
  22. Species Doris auriculata O. F. Müller, 1776 accepted as Facelina auriculata (O. F. Müller, 1776) (unaccepted > superseded combination)
  23. Species Doris barvicensis G. Johnston, 1838 accepted as Goniodoris nodosa (Montagu, 1808) accepted as Okenia nodosa (Montagu, 1808) (synonym)
  24. Species Doris beaumonti Farran, 1903 accepted as Diaphorodoris luteocincta (M. Sars, 1870) (synonym)
  25. Species Doris bifida A. E. Verrill, 1870 accepted as Acanthodoris pilosa (Abildgaard, 1789) (unaccepted > junior subjective synonym)
  26. Species Doris bifida Montagu, 1816 accepted as Hermaea bifida (Montagu, 1816) (original combination)
  27. Species Doris bilamellata Linnaeus, 1767 accepted as Onchidoris bilamellata (Linnaeus, 1767) (original combination)
  28. Species Doris biscayensis P. Fischer, 1872 accepted as Doris verrucosa Linnaeus, 1758
  29. Species Doris bodoensis Gunnerus, 1770 accepted as Aeolidia papillosa (Linnaeus, 1761) (synonym)
  30. Species Doris branchialis Rathke, 1806 accepted as Favorinus branchialis (Rathke, 1806)
  31. Species Doris caerulea Montagu, 1804 accepted as Tenellia caerulea (Montagu, 1804) accepted as Trinchesia caerulea (Montagu, 1804) (unaccepted > superseded combination)
  32. Species Doris caerulea Risso, 1826 accepted as Felimare villafranca (Risso, 1818) (synonym)
  33. Species Doris calcarae Vérany, 1846 accepted as Felimare picta (R. A. Philippi, 1836)
  34. Species Doris canariensis A. d'Orbigny, 1839 accepted as Platydoris argo (Linnaeus, 1767) (synonym)
  35. Species Doris clavigera O. F. Müller, 1776 accepted as Limacia clavigera (O. F. Müller, 1776) (original combination)
  36. Species Doris coccinea Forbes, 1848 accepted as Rostanga rubra (Risso, 1818)
  37. Species Doris complanata A. E. Verrill, 1880 accepted as Geitodoris planata (Alder & Hancock, 1846) (unaccepted > junior subjective synonym)
  38. Species Doris cornuta Rathke, 1806 accepted as Polycera quadrilineata (O. F. Müller, 1776)
  39. Species Doris coronata Gmelin, 1791 accepted as Doto coronata (Gmelin, 1791) (original combination)
  40. Species Doris depressa Alder & Hancock, 1842 accepted as Idaliadoris depressa (Alder & Hancock, 1842) (unaccepted > superseded combination)
  41. Species Doris derelicta P. Fischer, 1867 accepted as Doris verrucosa Linnaeus, 1758
  42. Species Doris diaphana Alder & Hancock, 1845 accepted as Onchidoris muricata (O. F. Müller, 1776) (synonym)
  43. Species Doris elegans Cantraine, 1835 accepted as Felimare picta (R. A. Philippi, 1836) (invalid: junior homonym of Doris elegans Quoy & Gaimard, 1832)
  44. Species Doris elegantula R. A. Philippi, 1844 accepted as Felimida elegantula (R. A. Philippi, 1844) (unaccepted > superseded combination)
  45. Species Doris elongata W. Thompson, 1840 accepted as Goniodoris nodosa (Montagu, 1808) accepted as Okenia nodosa (Montagu, 1808) (synonym)
  46. Species Doris eubalia P. Fischer, 1872 accepted as Doris sticta (Iredale & O'Donoghue, 1923) (unaccepted > junior subjective synonym)
  47. Species Doris fasciculata Gmelin, 1791 accepted as Fiona pinnata (Eschscholtz, 1831) (ICZN opinion 773: placed on the official index of rejected names)
  48. Species Doris flava Montagu, 1804 accepted as Polycera quadrilineata (O. F. Müller, 1776)
  49. Species Doris flavipes Leuckart, 1828 accepted as Doris pseudoargus Rapp, 1827 accepted as Archidoris pseudoargus (Rapp, 1827)
  50. Species Doris fumata Rüppell & Leuckart, 1830 accepted as Dendrodoris fumata (Rüppell & Leuckart, 1830) (original combination)
  51. Species Doris fusca O. F. Müller, 1776 accepted as Onchidoris bilamellata (Linnaeus, 1767)
  52. Species Doris glabra Friele & A. Hansen, 1876 accepted as Cadlina glabra (Friele & A. Hansen, 1876) (original combination)
  53. Species Doris gracilis Rapp, 1827 accepted as Felimare villafranca (Risso, 1818)
  54. Species Doris grandiflora Rapp, 1827 accepted as Dendrodoris grandiflora (Rapp, 1827)
  55. Species Doris guttata Risso, 1826 accepted as Dendrodoris grandiflora (Rapp, 1827) (synonym)
  56. Species Doris illuminata A. A. Gould, 1841 accepted as Polycera lessonii A. d'Orbigny, 1837 accepted as Palio dubia (M. Sars, 1829) (junior subjective synonym)
  57. Species Doris inconspicua Alder & Hancock, 1851 accepted as Atalodoris inconspicua (Alder & Hancock, 1851) (original combination)
  58. Species Doris infranaevata Abraham, 1877 accepted as Platydoris argo (Linnaeus, 1767) (dubious synonym)
  59. Species Doris infucata Rüppell & Leuckart, 1830 accepted as Hypselodoris infucata (Rüppell & Leuckart, 1830) (original combination)
  60. Species Doris johnstoni Alder & Hancock, 1845 accepted as Jorunna tomentosa (Cuvier, 1804)
  61. Species Doris krohni Vérany, 1846 accepted as Felimida krohni (Vérany, 1846) accepted as Rudmania krohni (Verany, 1846) (original combination)
  62. Species Doris lacera Abildgaard, 1806 accepted as Issena abildgaardi Pruvot-Fol, 1935 accepted as Colga villosa (Odhner, 1907) (preoccupied by Doris lacera Cuvier, 1804)
  63. Species Doris laevis Linnaeus, 1767 accepted as Cadlina laevis (Linnaeus, 1767) (unaccepted > superseded combination)
  64. Species Doris laevis J. E. Gray, 1850 accepted as Acanthodoris pilosa (Abildgaard, 1789) (synonym)
  65. Species Doris leuckartii Delle Chiaje, 1841 accepted as Doris pseudoargus Rapp, 1827 accepted as Archidoris pseudoargus (Rapp, 1827) (synonym)
  66. Species Doris limbata Cuvier, 1804 accepted as Dendrodoris limbata (Cuvier, 1804)
  67. Species Doris longicornis Montagu, 1808 accepted as Facelina auriculata (O. F. Müller, 1776) (dubious synonym)
  68. Species Doris loveni Alder & Hancock, 1862 accepted as Adalaria loveni (Alder & Hancock, 1862) (original combination)
  69. Species Doris lugubris Ehrenberg, 1828 accepted as Dendrodoris limbata (Cuvier, 1804) (unaccepted > junior subjective synonym)
  70. Species Doris luteocincta M. Sars, 1870 accepted as Diaphorodoris luteocincta (M. Sars, 1870) (original combination)
  71. Species Doris luteorosea Rapp, 1827 accepted as Felimida luteorosea (Rapp, 1827) (original combination)
  72. Species Doris lutescens Delle Chiaje, 1841 accepted as Felimare picta (R. A. Philippi, 1836) (dubious synonym)
  73. Species Doris maculata Garstang, 1896 accepted as Doris sticta (Iredale & O'Donoghue, 1923) (invalid: junior homonym of Doris maculata Montagu, 1804)
  74. Species Doris maura Forbes, 1840 accepted as Aegires punctilucens (A. d'Orbigny, 1837)
  75. Species Doris millegrana Alder & Hancock, 1854 accepted as Aporodoris millegrana (Alder & Hancock, 1854) (original combination)
  76. Species Doris muricata O. F. Müller, 1776 accepted as Onchidoris muricata (O. F. Müller, 1776) (original combination)
  77. Species Doris nardii Vérany, 1846 accepted as Felimare picta (R. A. Philippi, 1836) (synonym)
  78. Species Doris nigricans J. Fleming, 1820 accepted as Acanthodoris pilosa (Abildgaard, 1789) (synonym)
  79. Species Doris nigricans Otto, 1823 accepted as Dendrodoris limbata (Cuvier, 1804) (synonym)
  80. Species Doris nodosa Montagu, 1808 accepted as Goniodoris nodosa (Montagu, 1808) accepted as Okenia nodosa (Montagu, 1808)
  81. Species Doris oblonga Alder & Hancock, 1845 accepted as Knoutsodonta oblonga (Alder & Hancock, 1845) accepted as Atalodoris oblonga (Alder & Hancock, 1845) (original combination)
  82. Species Doris obsoleta Rüppell & Leuckart, 1830 accepted as Goniobranchus obsoletus (Rüppell & Leuckart, 1830) (original combination)
  83. Species Doris obvelata O. F. Müller, 1776 accepted as Cadlina laevis (Linnaeus, 1767) (synonym)
  84. Species Doris orbignyi H. Adams & A. Adams, 1858 accepted as “Montereina” punctifera (Abraham, 1877) represented as Peltodoris punctifera (Abraham, 1877) (Invalid: junior homonym of Doris orbignyi Férussac, 1822; Doris punctifera is a replacement name)
  85. Species Doris orsinii Vérany, 1846 accepted as Felimare orsinii (Vérany, 1846) (original combination)
  86. Species Doris pareti Vérany, 1846 accepted as Goniodoris castanea Alder & Hancock, 1845 accepted as Pelagella castanea (Alder & Hancock, 1845) (unaccepted > junior subjective synonym)
  87. Species Doris pasini Vérany, 1846 accepted as Felimare villafranca (Risso, 1818)
  88. Species Doris pedata Montagu, 1816 accepted as Edmundsella pedata (Montagu, 1816)
  89. Species Doris pellucida Risso, 1826 accepted as Cadlina pellucida (Risso, 1826)
  90. Species Doris pennigera Montagu, 1813 accepted as Thecacera pennigera (Montagu, 1813)
  91. Species Doris peregrina Gmelin, 1791 accepted as Cratena peregrina (Gmelin, 1791) (original combination)
  92. Species Doris philippii Weinkauff, 1873 accepted as Jorunna tomentosa (Cuvier, 1804) (dubious synonym)
  93. Species Doris picta R. A. Philippi, 1836 accepted as Felimare picta (R. A. Philippi, 1836) (unaccepted > superseded combination)
  94. Species Doris pilosa Abildgaard, 1789 accepted as Acanthodoris pilosa (Abildgaard, 1789)
  95. Species Doris pinnatifida Montagu, 1804 accepted as Doto pinnatifida (Montagu, 1804)
  96. Species Doris piraini Vérany, 1846 accepted as Felimida purpurea (Risso, 1831) accepted as Rudmania purpurea (Risso, 1831)
  97. Species Doris planata Alder & Hancock, 1846 accepted as Geitodoris planata (Alder & Hancock, 1846) (original combination)
  98. Species Doris planulata W. Stimpson, 1853 accepted as Cadlina laevis (Linnaeus, 1767) (unaccepted > junior subjective synonym, dubious synonym)
  99. Species Doris porri Vérany, 1846 accepted as Paradoris indecora (Bergh, 1881) (synonym)
  100. Species Doris proxima Alder & Hancock, 1854 accepted as Adalaria proxima (Alder & Hancock, 1854) (original combination)
  101. Species Doris pseudoargus Rapp, 1827 accepted as Archidoris pseudoargus (Rapp, 1827) (unaccepted > superseded combination)
  102. Species Doris pulchella Aradas, 1847 accepted as Doris bicolor (Bergh, 1884) (primary homonym of Doris pulchella Rüppell & Leuckart, 1828)
  103. Species Doris pulcherrima Cantraine, 1835 accepted as Felimare villafranca (Risso, 1818)
  104. Species Doris punctata A. d'Orbigny, 1839 accepted as “Montereina” punctifera (Abraham, 1877) represented as Peltodoris punctifera (Abraham, 1877) (Invalid: junior homonym of Doris punctata Rüppell & Leuckart, 1831, and D. punctata Quoy & Gaimard, 1832; Doris orbignyi H. & A. Adams, 1858, is a replacement name)
  105. Species Doris punctifera Abraham, 1877 accepted as “Montereina” punctifera (Abraham, 1877) represented as Peltodoris punctifera (Abraham, 1877) (original combination)
  106. Species Doris purpurea Risso, 1831 accepted as Felimida purpurea (Risso, 1831) accepted as Rudmania purpurea (Risso, 1831) (original combination)
  107. Species Doris pusilla Alder & Hancock, 1845 accepted as Atalodoris pusilla (Alder & Hancock, 1845) (original combination)
  108. Species Doris quadrangulata Jeffreys, 1869 accepted as Acanthodoris pilosa (Abildgaard, 1789) (synonym)
  109. Species Doris quadricolor Rüppell & Leuckart, 1830 accepted as Chromodoris quadricolor (Rüppell & Leuckart, 1830) (original combination)
  110. Species Doris quadricornis Montagu, 1813 accepted as Okenia aspersa (Alder & Hancock, 1845) (ICZN opinion 1014: suppressed under the plenary powers for the purpose of the Principle of Priority)
  111. Species Doris quadrilineata O. F. Müller, 1776 accepted as Polycera quadrilineata (O. F. Müller, 1776)
  112. Species Doris radiata Gmelin, 1791 accepted as Glaucus atlanticus Forster, 1777 (synonym)
  113. Species Doris ramosa Cantraine, 1835 accepted as Kaloplocamus ramosus (Cantraine, 1835)
  114. Species Doris rappii Cantraine, 1841 (unaccepted, synonym)
  115. Species Doris repanda Alder & Hancock, 1842 (unaccepted)
  116. Species Doris reticulata R. A. Philippi, 1836 (unaccepted > junior homonym, junior homonym of Doris reticulata Quoy & Gaimard, 1832)
  117. Species Doris rocinela Leach, 1852 (unaccepted, synonym)
  118. Species Doris rubra Risso, 1818 (unaccepted, original combination)
  119. Species Doris scacchi Delle Chiaje, 1830 (unaccepted)
  120. Species Doris schembrii Vérany, 1846 (unaccepted)
  121. Species Doris schultzii Delle Chiaje, 1841 (unaccepted)
  122. Species Doris setigera Rapp, 1827 (unaccepted, synonym)
  123. Species Doris sibirica Aurivillius, 1887 (unaccepted)
  124. Species Doris similis Alder & Hancock, 1842 (unaccepted, synonym)
  125. Species Doris sismondae Vérany, 1846 (unaccepted, synonym)
  126. Species Doris sparsa Alder & Hancock, 1846 (unaccepted, original combination)
  127. Species Doris stellata Gmelin, 1791 (unaccepted, synonym)
  128. Species Doris sublaevis W. Thompson, 1840 (unaccepted, synonym)
  129. Species Doris subquadrata Alder & Hancock, 1845 (unaccepted, synonym)
  130. Species Doris subtumida Abraham, 1877 (unaccepted, synonym)
  131. Species Doris tenera O. G. Costa, 1840 (unaccepted)
  132. Species Doris testudinaria Risso, 1826 (unaccepted, dubious synonym)
  133. Species Doris tomentosa Cuvier, 1804 (unaccepted, original combination)
  134. Species Doris tricolor Cantraine, 1835 (unaccepted, original combination)
  135. Species Doris ulidiana W. Thompson, 1845 (unaccepted, synonym)
  136. Species Doris valenciennesi Cantraine, 1841 (unaccepted)
  137. Species Doris vermigera W. Turton, 1807 (unaccepted > junior subjective synonym)
  138. Species Doris villae Vérany, 1846 (unaccepted)
  139. Species Doris villafranca Risso, 1818 (unaccepted, original combination)
  140. Species Doris virescens Risso, 1826 (unaccepted, synonym)
  141. Species Doris zetlandica Alder & Hancock, 1854 (unaccepted > superseded combination)
marine
Linnaeus, C. (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii. Holmiae [Stockholm]. [iii] + 824 pp. , available online at https://biodiversitylibrary.org/page/726886
page(s): 653 [details] Available for editors  PDF available
MolluscaBase eds. (2025). MolluscaBase. Doris Linnaeus, 1758. Accessed through: Costello, M.J.; Bouchet, P.; Boxshall, G.; Arvanitidis, C.; Appeltans, W. (2025) European Register of Marine Species at: https://www.marbef.org/data/aphia.php?p=taxdetails&id=137914 on 2026-07-29
Costello, M.J.; Bouchet, P.; Boxshall, G.; Arvanitidis, C.; Appeltans, W. (2026). European Register of Marine Species. Doris Linnaeus, 1758. Accessed at: https://www.vliz.be/vmdcdata/narms/narms.php?p=taxdetails&id=137914 on 2026-07-29
Date
action
by
2004-12-21 15:54:05Z
created
Gofas, Serge
2013-06-25 22:58:08Z
changed
Bouchet, Philippe
2015-03-27 11:07:58Z
changed
Gofas, Serge

original description Linnaeus, C. (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii. Holmiae [Stockholm]. [iii] + 824 pp. , available online at https://biodiversitylibrary.org/page/726886
page(s): 653 [details] Available for editors  PDF available

original description (of Staurodoris Bergh, 1878) Bergh, L. S. R. (1878). Malacologische Untersuchungen. In: C. Semper (ed.) Reisen im Archipel der Philippinen, Theil 2, Wissenschaftliche Resultate, Band 2, Theil 2, Heft 13: 547-601, pls 62-65; Heft 14: 603-645, I-L, pls 66-68. , available online at https://biodiversitylibrary.org/page/14374143 [details] 

original description (of Doridigitata A. d'Orbigny, 1839) Orbigny, A. d'. (1836-1842). <i>Mollusques, Echinodermes, Foraminifčres et Polypiers recueillis aux Iles Canaries par MM. Webb et Berthelot</i>. In: Barker-Webb, P. & Berthelot, S. (eds) <i>Histoire naturelle des Iles Canaries</i>, volume 2(2): 117 pp., pls 1-7, 7B. pp. 1-24 [Aug. 1839], 25-48 [Sept. 1839], 49-72 [Oct-1839], 73-104 [Jan. 1840], 105-136 [Mar. 1840], 137-143 [Apr. 1840], 145-152 [Aug. 1842]; pl. 1 [Jul. 1836], 2 [Dec. 1836], 3 [May 1842], 4-5 [June 1840], 7 [May 1842], 6, 7B [Aug. 1842]. Paris: Béthune. In: P. B. Webb & S. Berthelot (1836-1844), Histoire naturelle des Iles Canaries 2(2). Zoologie: Mollusques. Paris., available online at https://www.biodiversitylibrary.org/page/41308382 [details] 

original description (of Siraius Er. Marcus, 1955) Marcus, Er. (1955). Opisthobranchia from Brazil. <em>Boletim da Faculdade de Filosofia, Cięncias e Letras da Universidade de Săo Paulo (Zoologia).</em> 20: 89-261., available online at https://www.revistas.usp.br/bffclzoologia/article/view/120213 [details] Available for editors  PDF available

original description (of Doris (Doridigitata) A. d'Orbigny, 1839) Orbigny, A. d'. (1836-1842). <i>Mollusques, Echinodermes, Foraminifčres et Polypiers recueillis aux Iles Canaries par MM. Webb et Berthelot</i>. In: Barker-Webb, P. & Berthelot, S. (eds) <i>Histoire naturelle des Iles Canaries</i>, volume 2(2): 117 pp., pls 1-7, 7B. pp. 1-24 [Aug. 1839], 25-48 [Sept. 1839], 49-72 [Oct-1839], 73-104 [Jan. 1840], 105-136 [Mar. 1840], 137-143 [Apr. 1840], 145-152 [Aug. 1842]; pl. 1 [Jul. 1836], 2 [Dec. 1836], 3 [May 1842], 4-5 [June 1840], 7 [May 1842], 6, 7B [Aug. 1842]. Paris: Béthune. In: P. B. Webb & S. Berthelot (1836-1844), Histoire naturelle des Iles Canaries 2(2). Zoologie: Mollusques. Paris., available online at https://www.biodiversitylibrary.org/page/41308382
page(s): 39, 40 [details] 

basis of record Valdés Á. (2002). A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia). <em>Zoological Journal of the Linnean Society.</em> 136: 535-636., available online at https://doi.org/10.1046/j.1096-3642.2002.00039.x
page(s): 540-560 [details] Available for editors  PDF available
 
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To European Nucleotide Archive, ENA (Doris) 
To ITIS
Unreviewed